This paleobotany list records new fossil plant taxa that were to be described during the year 2012, as well as notes other significant paleobotany discoveries and events which occurred during 2012.

List of years in paleobotany
In paleontology
2009
2010
2011
2012
2013
2014
2015
In arthropod paleontology
2009
2010
2011
2012
2013
2014
2015
In paleoentomology
2009
2010
2011
2012
2013
2014
2015
In paleomalacology
2009
2010
2011
2012
2013
2014
2015
In reptile paleontology
2009
2010
2011
2012
2013
2014
2015
In archosaur paleontology
2009
2010
2011
2012
2013
2014
2015
In mammal paleontology
2009
2010
2011
2012
2013
2014
2015
In paleoichthyology
2009
2010
2011
2012
2013
2014
2015
List of years in paleobotany
In paleontology
2009
2010
2011
2012
2013
2014
2015
In arthropod paleontology
2009
2010
2011
2012
2013
2014
2015
In paleoentomology
2009
2010
2011
2012
2013
2014
2015
In paleomalacology
2009
2010
2011
2012
2013
2014
2015
In reptile paleontology
2009
2010
2011
2012
2013
2014
2015
In archosaur paleontology
2009
2010
2011
2012
2013
2014
2015
In mammal paleontology
2009
2010
2011
2012
2013
2014
2015
In paleoichthyology
2009
2010
2011
2012
2013
2014
2015

Bryophytes

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Name Novelty Status Authors Age Type locality Location Notes Images

Bryokhutuliinia crassimarginata[1]

Sp nov

Ignatov et al

Late Jurassic

Ukurey formation

  Russia

A moss of uncertain placement.

Lycophytes

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Name Novelty Status Authors Age Type locality Location Notes Images

Winslowia[2]

Gen. et sp. nov

Valid

Dunn et al.

Carboniferous (Serpukhovian)

Pride Mountain Formation

  United States

A member of Isoetales belonging to the group Chaloneriaceae. Genus includes new species W. tuscumbiana.

Marchantiophytes

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Name Novelty Status Authors Age Type locality Location Notes Images

Riccardiothallus[3]

Gen. et sp. nov

Valid

Guo et al.

Devonian (Pragian)

Posongchong Formation

  China

A liverwort belonging to the family Aneuraceae. Genus includes new species R. devonicus.

Ferns and fern allies

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Name Novelty Status Authors Age Type locality Location Notes Images

Korallipteris[4]

Gen. et comb. nov

Valid

Vera & Passalia

Triassic to Miocene

Antarctica
  Argentina
  Chile
  New Zealand[5]

A fern of uncertain phylogenetic placement.
A new genus for "Gleichenia" argentinica (1962);
Additional species were originally placed in Gleichenia, Gleichenites and Microphyllopteris.

Millerocaulis tekelili[6]

Sp. nov

Valid

Vera

Early Cretaceous (Aptian)

Cerro Negro Formation

Antarctica
(Livingston Island)

An Osmundaceae fern. Originally placed in Millerocaulis;
Moved to Claytosmunda tekelili in (2017).[7]

Osmundopsis rafaelii[8]

Sp. nov

Valid

Escapa & Cúneo

Early Jurassic

  Argentina

A member of Osmundaceae.

Todites cacereii[8]

Sp. nov

Valid

Escapa & Cúneo

Early Jurassic

  Argentina

A member of Osmundaceae.

Bennettitales

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Name Novelty Status Authors Age Type locality Location Notes Images

Anomozamites harrisi[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Anomozamites heeri[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Anomozamites villosus[10]

Sp. nov

Valid

Pott et al.

Probably late Middle Jurassic

Daohugou Beds

  China

A member of Bennettitales. Originally described as a species of Anomozamites
Moved to the Wielandiella villosus (2015).[11][12]

Cycadolepis barnardii[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Cycadolepis karatauensis[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Cycadolepis kirillii[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Cycadolepis kokalensis[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Cycadolepis lata[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Cycadolepis maija[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Cycadolepis multistomata[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Cycadolepis pachydermatica[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Cycadolepis perfecta[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Cycadolepis prynadае[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Cycadolepis schairensis[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Nilssoniopteris chirchilica[9]

Sp. nov

Valid

Kiritchkova & Nosova

Middle Jurassic (Aalenian)

Кokala Suite

  Kazakhstan

Nilssoniopteris localis[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Nilssoniopteris saveljevii[9]

Sp. nov

Valid

Kiritchkova & Nosova

Middle Jurassic (Aalenian)

Tonasha Formation

  Kazakhstan

Pterophyllum chagaense[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Pterophyllum doludenkoae[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Pterophyllum originalum[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Pterophyllum pаrvisegmentatum[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Pterophyllum pinniforme[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Pterophyllum valentinaeа[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Pterophyllum vsevolodii[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Williamsonia antonia[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Williamsonia baranovaeа[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Williamsonia chagaensе[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Williamsonia kokalensis[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Williamsonia vialovaea[9]

Sp. nov

Valid

Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

"Pteridospermatophytes"

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Name Novelty Status Authors Age Type locality Location Notes Images

Ctenozamites schweitzerii[9]

Sp. nov

Valid

Kiritchkova & Nosova

Middle Jurassic (Bajocian)

Karadiirmen' Formation

  Kazakhstan

A member of Pteridospermatophyta of uncertain affinities.

Eretmonia macloughlinii[13]

Sp. nov

Valid

Ryberg, Taylor & Taylor

Late Permian

Upper Buckley Formation

  Antarctica

A member of Glossopteridales.

Kuvakospermum[14]

Gen. et sp. nov

Valid

Naugolnykh & Sidorov

Permian (Wordian)

  Russia

A seed fern belonging to the family Peltaspermaceae. Genus includes new species K. pedatum.

Lidgettoniopsis[15]

Gen. et sp. nov

Valid

Ryberg, Taylor & Taylor

Late Permian

Beacon Supergroup (upper Buckley Formation)

  Antarctica

A seed fern belonging to the group Dictyopteridiales and the family Eretmoniaceae. Genus includes new species L. ramulus.

Scutum leiophyllum[15]

Sp. nov

Valid

Ryberg, Taylor & Taylor

Late Permian

Beacon Supergroup (upper Buckley Formation)

  Antarctica

A member of Glossopteridales belonging to the family Dictyopteridiaceae.

Vetlugospermum[16]

Gen. et sp. nov

Valid

Naugolnykh

Early Triassic

  Russia

A seed fern belonging to the group Peltaspermales. Genus includes new species V. rombicum.

Ginkgoales

edit
Name Novelty Status Authors Age Type locality Location Notes Images

Baiera kazachstanica[9]

Sp. nov

Valid

Nosova & Kiritchkova in Kiritchkova & Nosova

Early Jurassic (Toarcian)

Кokala Suite

  Kazakhstan

Baieroxylon rocablanquense[17]

Sp. nov

Valid

Gnaedinger

Early Jurassic

Roca Blanca Formation

  Argentina

Ginkgo cranei[18]

Sp. nov

Valid

Zhou, Quan, & Liu

upper Paleocene

Sentinel Butte Formation

  USA
  North Dakota

An extinct species in the modern genus Ginkgo

Ginkgo gomolitzkyana[19]

Sp. nov

Valid

Nosova

Middle Jurassic

Angren Formation

  Uzbekistan

A member of Ginkgoales.

Ginkgo huolinhensis[20]

Sp. nov

Valid

Dong & Sun

Early Cretaceous

Huolinhe Formation

  China

Ginkgoites barnardianus[9]

Sp. nov

Valid

Nosova & Kiritchkova in Kiritchkova & Nosova

Middle Jurassic (Aalenian)

Кokala Suite

  Kazakhstan

Ginkgoites irdalicus[9]

Sp. nov

Valid

Nosova & Kiritchkova in Kiritchkova & Nosova

Early Jurassic

Ergozi Suite

  Kazakhstan

Ginkgoites kelendensis[9]

Sp. nov

Valid

Nosova & Kiritchkova in Kiritchkova & Nosova

Middle Jurassic (Aalenian)

Tonasha Formation

  Kazakhstan

Ginkgoites ketovianus[9]

Sp. nov

Valid

Nosova & Kiritchkova in Kiritchkova & Nosova

Middle Jurassic (Bathonian)

Sarydiirmeny Suite

  Kazakhstan

Leptotoma asiatica[9]

Sp. nov

Valid

Nosova & Kiritchkova in Kiritchkova & Nosova

Middle Jurassic (Bajocian)

Karadiirmen' Formation

  Kazakhstan

Pseudotorellia karadiirmenica[9]

Sp. nov

Valid

Nosova & Kiritchkova in Kiritchkova & Nosova

Middle Jurassic (Bajocian)

Karadiirmen' Formation

  Kazakhstan

Conifers

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Araucariaceae

edit
Name Novelty Status Authors Age Unit Location Notes Images

Emwadea[21]

Gen. et sp. nov

Valid

Dettmann, Clifford & Peters

Early Cretaceous (Albian)

Winton Formation

  Australia

A member of the family Araucariaceae. Genus includes new species E. microcarpa.

Cheirolepidiaceae

edit
Name Novelty Status Authors Age Unit Location Notes Images

Duartenia[22]

Gen. et sp. nov

Valid

Mohr et al.

Early Cretaceous

Crato Formation

  Brazil

A possible member of Cheirolepidiaceae. The type species is Duartenia araripensis.

Glenrosa falcata[23]

Sp. nov

Valid

Gomez, Ewin & Daviero-Gomez

Early Cretaceous (Barremian)

Calizas de La Huergina Formation

  Spain

A conifer.

Cupressaceae

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Name Novelty Status Authors Age Unit Location Notes Images

Calocedrus huashanensis[24]

Sp. nov

Valid

Shi, Zhou, & Xie

Oligocene

Ningming Formation

  China

A Calocedrus species

Fokienia shengxianensis[25]

Sp. nov

Valid

He, Sun & Liu

Late Miocene

Shengxian Formation
Xiaolongtan Formation

  China

Originally placed in Fokienia.
Moved to Calocedrus shengxianensis in (2015).[26]

Pinaceae

edit
Name Novelty Status Authors Age Unit Location Notes Images

Eathiestrobus[27]

Gen. et sp. nov

Valid

Rothwell et al.

Late Jurassic (Kimmeridgian)

Kimmeridge Clay Formation

  United Kingdom

A member of Pinaceae. The type species is Eathiestrobus mackenziei.

Picea burtonii[28]

Sp. nov

Valid

Klymiuk & Stockey

Early Cretaceous (Valanginian)

  Canada

A spruce species seed cone.

Pinus yorkshirensis[29]

Sp. nov

Valid

Ryberg et al.

Early Cretaceous (Hauterivian-Barremian)

Speeton Clay Formation

  United Kingdom

A pine.

Podocarpaceae

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Name Novelty Status Authors Age Unit Location Notes Images

Dacrycarpus puertae[30]

Sp. nov

Valid

Wilf

Eocene

  Argentina

A species of Dacrycarpus

Taxaceae

edit
Name Novelty Status Authors Age Unit Location Notes Images

Marskea kazachstanica[9]

Sp. nov

Valid

Nosova & Kiritchkova in Kiritchkova & Nosova

Early Jurassic (Toarcian)

Kokala Suite

  Kazakhstan

Torreya kazachstanica[9]

Sp. nov

Valid

Nosova & Kiritchkova in Kiritchkova & Nosova

Early Jurassic (Toarcian)

Kokala Suite

  Kazakhstan

Other conifers

edit
Name Novelty Status Authors Age Unit Location Notes Images

Elatocladus bulakus[9]

Sp. nov

Valid

Nosova & Kiritchkova in Kiritchkova & Nosova

Early Jurassic (Toarcian)

Kokala Suite

  Kazakhstan

A conifer of uncertain affinities.

Plyophyllioxylon[31]

Gen. et sp. nov

Valid

Feng et al.

Permian (Roadian/Wordian)

Lower Shihhotse Formation

  China

A conifer of uncertain phylogenetic placement. Genus includes new species P. hulstaiense.

Other gymnosperms

edit
Name Novelty Status Authors Age Type locality Location Notes Images

Nilssonia prosvirjakovaea[9]

Sp. nov

Valid

Kiritchkova & Nosova

Middle Jurassic (Bajocian)

Karadiirmen' Formation

  Kazakhstan

A cycad.

Rehezamites[32]

Gen. et sp. nov

Valid

Pott et al.

Early Cretaceous (Aptian)

Yixian Formation

  China

A seed plant of uncertain phylogenetic placement. Genus includes new species R. anisolobus.

Angiosperms

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Name Novelty Status Authors Age Unit Location Notes Images

Archaeampelos senonica[33]

Sp. nov

Valid

Alekseev

Late Cretaceous

  Russia

Baasia[34]

Gen. et sp. nov

Valid

Estrada-Ruiz et al.

Late Cretaceous (probably early to middle Maastrichtian)

McRae Formation

  United States

A member of the family Celastraceae. Genus includes new species B. armendarisense.

Bertilanthus[35]

Gen. et sp. nov

Valid

Friis & Pedersen

Late Cretaceous

  Sweden

An asterid. Genus includes new species B. scanicus.

Camptodromites sibiricus[33]

Sp. nov

Valid

Alekseev

Late Cretaceous

  Russia

Celastrinites lanceolatus[33]

Sp. nov

Valid

Alekseev

Late Cretaceous

  Russia

Chaneya hainanensis[36]

Sp. nov

Valid

Feng & Jin

Eocene

Changchang Basin

  China

A fossil fruit.

Cissites basicordatus[33]

Sp. nov

Valid

Alekseev

Late Cretaceous

  Russia

Cissus lombardii[37]

Sp. nov

Valid

Manchester, Chen & Lott

Early Oligocene

Mancora Formation

  Peru

A species of Cissus.

Eucalyptus caldericola[38]

Sp. nov

Valid

Hermsen, Gandolfo & Zamaloa

Eocene (Ypresian)

  Argentina

A species of Eucalyptus.

Eucalyptus frenguelliana[38]

Sp. nov

Valid

Gandolfo & Zamaloa in Hermsen, Gandolfo & Zamaloa

Eocene (Ypresian)

  Argentina

A species of Eucalyptus.

Eucalyptus lynchiae[38]

Sp. nov

Valid

Gandolfo & Hermsen in Hermsen, Gandolfo & Zamaloa

Eocene (Ypresian)

  Argentina

A species of Eucalyptus.

Fulleroxylon[34]

Gen. et sp. nov

Valid

Estrada-Ruiz et al.

Late Cretaceous (probably early to middle Maastrichtian)

McRae Formation

  United States

A member of the family Myrtaceae. Genus includes new species F. armendarisense.

Juglandiphyllites microdentatus[33]

Sp. nov

Valid

Alekseev

Late Cretaceous

  Russia

Laurinoxylon stichkai[39]

Sp. nov

Valid

Boonchai & Manchester

Early Eocene

Bridger Formation

  United States

A member of Lauraceae described on the basis of fossil wood.

Laurophyllum calicarioides[40]

Sp. nov

Valid

Bannister, Conran & Lee

Early Miocene

  New Zealand

A member of Lauraceae with affinities to members of the genus Litsea.

Laurophyllum lacustris[40]

Sp. nov

Valid

Bannister, Conran & Lee

Early Miocene

  New Zealand

A member of Lauraceae with affinities to members of the genus Beilschmiedia.

Laurophyllum maarensis[40]

Sp. nov

Valid

Bannister, Conran & Lee

Early Miocene

  New Zealand

A member of Lauraceae with affinities to members of the genus Cryptocarya.

Laurophyllum microphyllum[40]

Sp. nov

Valid

Bannister, Conran & Lee

Early Miocene

  New Zealand

A member of Lauraceae with affinities to members of the genus Cryptocarya.

Laurophyllum otagoensis[40]

Sp. nov

Valid

Bannister, Conran & Lee

Early Miocene

  New Zealand

A member of Lauraceae with affinities to members of the genus Beilschmiedia.

Laurophyllum sylvestris[40]

Sp. nov

Valid

Bannister, Conran & Lee

Early Miocene

  New Zealand

A member of Lauraceae with affinities to members of the genus Beilschmiedia.

Laurophyllum taieriensis[40]

Sp. nov

Valid

Bannister, Conran & Lee

Early Miocene

  New Zealand

A member of Lauraceae with affinities to members of the genus Cryptocarya.

Laurophyllum vulcanicola[40]

Sp. nov

Valid

Bannister, Conran & Lee

Early Miocene

  New Zealand

A member of Lauraceae with affinities to members of the genus Beilschmiedia.

Laurophyllum waipiata[40]

Sp. nov

Valid

Bannister, Conran & Lee

Early Miocene

  New Zealand

A member of Lauraceae with affinities to members of the genus Cryptocarya.

Litseopsis nova-zelandiae[40]

Sp. nov

Valid

Bannister, Conran & Lee

Early Miocene

  New Zealand

A member of Lauraceae.

Mahonia grimmii[41]

Sp. nov

Valid

Güner & Denk

Miocene

Eskihisar Formation

  Turkey

A species of Mahonia.

Mahonia somaensis[41]

Sp. nov

Valid

Güner & Denk

Miocene

Soma Formation

  Turkey

A species of Mahonia.

Morinda chinensis[42]

Sp. nov

Valid

Shi et al.

Eocene

Changchang Formation

  China

A species of Morinda.

Newtonia mushensis[43]

Sp. nov

Valid

Pan et al.

Early Miocene

  Ethiopia

A species of Newtonia.

Palaeophytocrene hammenii[44]

Sp. nov

Valid

Stull in Stull et al.

Middle-late Paleocene

Bogotá Formation

  Colombia

A member of Icacinaceae belonging to the tribe Phytocreneae.

Palaeophytocrene piggae[44]

Sp. nov

Valid

Stull in Stull et al.

Late Paleocene

Sentinel Butte Formation

  United States

A member of Icacinaceae belonging to the tribe Phytocreneae.

Pygmaeoxylon[34]

Gen. et sp. nov

Valid

Estrada-Ruiz et al.

Late Cretaceous (probably early to middle Maastrichtian)

McRae Formation

  United States

A magnoliid of uncertain phylogenetic placement. Genus includes new species P. paucipora.

Pyrenacantha austroamericana[44]

Sp. nov

Valid

Stull in Stull et al.

Late early Oligocene

  Peru

A species of Pyrenacantha.

Schenkeriphyllum glanduliferum[45]

Gen. et sp. nov

Valid

Mohr, Coiffard & Bernardes-de-Oliveira

Early Cretaceous (Aptian)

Crato Formation

  Brazil

Fossil magnolialean angiosperm.

Wilsonoxylon[39]

Gen. et sp. nov

Valid

Boonchai & Manchester

Early Eocene

Bridger Formation

  United States

A member of Canellaceae described on the basis of fossil wood. Genus includes new species W. edenense.

References

edit
  1. ^ Ignatov, M. S.; Karasev, E. V.; Sinitsa, S. M.; Maslova, E. V. (2013). "New Bryokhutuliinia species (Bryophyta) with sporophytes from the Upper Jurassic of Transbaikalia". Arctoa. 22: 69–78. doi:10.15298/arctoa.22.10.
  2. ^ Michael T. Dunn; Prescott Atkinson; James Lacefield; Michael Rischbieter (2012). "Winslowia tuscumbiana gen. et sp. nov. (Chaloneriaceae): A Cormose, Heterosporous, Ligulate Lycopsid Reconstructed from the Inside Out from the Pride Mountain Formation (Late Mississippian/Serpukhovian) of Northern Alabama". International Journal of Plant Sciences. 173 (1): 96–111. doi:10.1086/662679. S2CID 85874193.
  3. ^ Cai-Qing Guo; Dianne Edwards; Peng-Cheng Wu; Jeffrey G. Duckett; Francis M. Hueber; Cheng-Sen Li (2012). "Riccardiothallus devonicus gen. et sp. nov., the earliest simple thalloid liverwort from the Lower Devonian of Yunnan, China". Review of Palaeobotany and Palynology. 176–177: 35–40. Bibcode:2012RPaPa.176...35G. doi:10.1016/j.revpalbo.2012.03.012.
  4. ^ Ezequiel I. Vera; Mauro G. Passalia (2012). "Korallipteris, a new genus for Mesozoic Gleichenia-like fern fronds". Geobios. 45 (4): 421–428. Bibcode:2012Geobi..45..421V. doi:10.1016/j.geobios.2011.09.002. hdl:11336/68155.
  5. ^ John G. Conran; Joseph A. Jackson; Daphne E. Lee; Elizabeth M. Kennedy (2017). "Gleichenia-like Korallipteris alineae sp. nov. macrofossils (Polypodiophyta) from the Miocene Landslip Hill silcrete, New Zealand". New Zealand Journal of Botany. 55 (3): 258–275. Bibcode:2017NZJB...55..258C. doi:10.1080/0028825X.2017.1317278. S2CID 91077403.
  6. ^ Ezequiel I. Vera (2012). "Millerocaulis tekelili sp. nov., a new species of osmundalean fern from the Aptian Cerro Negro Formation (Antarctica)". Alcheringa: An Australasian Journal of Palaeontology. 36 (1): 35–45. Bibcode:2012Alch...36...35V. doi:10.1080/03115518.2011.576541. S2CID 129810694.
  7. ^ Benjamin Bomfleur; Guido W. Grimm; Stephen McLoughlin (2017). "The fossil Osmundales (Royal Ferns)—a phylogenetic network analysis, revised taxonomy, and evolutionary classification of anatomically preserved trunks and rhizomes". PeerJ. 5: e3433. doi:10.7717/peerj.3433. PMC 5508817. PMID 28713650.
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