2015 in reptile paleontology

This list of fossil reptiles described in 2015 is a list of new taxa of fossil reptiles that were described during the year 2015, as well as other significant discoveries and events related to reptile paleontology that occurred in 2015.

List of years in reptile paleontology
In archosaur paleontology
2012
2013
2014
2015
2016
2017
2018
In paleobotany
2012
2013
2014
2015
2016
2017
2018
In paleontology
2012
2013
2014
2015
2016
2017
2018
In arthropod paleontology
2012
2013
2014
2015
2016
2017
2018
In paleoentomology
2012
2013
2014
2015
2016
2017
2018
In paleomalacology
2012
2013
2014
2015
2016
2017
2018
In mammal paleontology
2012
2013
2014
2015
2016
2017
2018
In paleoichthyology
2012
2013
2014
2015
2016
2017
2018
In science
2012
2013
2014
2015
2016
2017
2018

Ichthyosauromorphs edit

Research edit

New taxa edit

Name Novelty Status Authors Age Unit Location Notes Images

Cartorhynchus[3]

Gen. et sp. nov

Valid

Motani et al.

Early Triassic (Olenekian)

Nanlinghu Formation

  China

A relative of ichthyopterygians. The type species is Cartorhynchus lenticarpus.

Dearcmhara[4]

Gen. et sp. nov

Valid

Brusatte et al.

Jurassic (Toarcian–Bajocian)

  United Kingdom

A basal member of Neoichthyosauria. The type species is Dearcmhara shawcrossi.

Eretmorhipis[5]

Gen. et sp. nov

Valid

Chen et al.

Early Triassic (late Spathian)

Jialingjiang Formation

  China

A hupehsuchian. The type species is Eretmorhipis carrolldongi.

Ichthyosaurus anningae[6]

Sp. nov

Valid

Lomax & Massare

Early Jurassic (Hettangian/Sinemurian–Pliensbachian)

  United Kingdom

A species of Ichthyosaurus.

Muiscasaurus[7]

Gen. et sp. nov

Valid

Maxwell et al.

Early Cretaceous (Barremian–Aptian)

Paja Formation

  Colombia

A member of Ophthalmosauridae. The type species is Muiscasaurus catheti.

Sauropterygians edit

Research edit

New taxa edit

Name Novelty Status Authors Age Unit Location Notes Images

Anguanax[11]

Gen. et sp. nov

Valid

Cau & Fanti

Late Jurassic (Oxfordian)

Rosso Ammonitico Veronese Formation

  Italy

A pliosaurid. The type species is Anguanax zignoi.

Atychodracon[12]

Gen. et comb. nov

Valid

Smith

Early Jurassic

  United Kingdom

A rhomaleosaurid plesiosaur; a new genus for "Plesiosaurus" megacephalus Stutchbury (1846).

Cardiocorax[13]

Gen. et sp. nov

Valid

Araújo et al.

Late Cretaceous (early Maastrichtian)

Mocuio Formation

  Angola

An elasmosaurid plesiosaur. The type species is Cardiocorax mukulu.

Dianmeisaurus[14]

Gen. et sp. nov

Valid

Shang & Li

Middle Triassic (Anisian)

Guangling Formation

  China

A member of Eosauropterygia of uncertain phylogenetic placement, more closely related to nothosaurs than to plesiosaurs. The type species is Dianmeisaurus gracilis.

Makhaira[15]

Gen. et sp. nov

Valid

Fischer et al.

Early Cretaceous (Hauterivian)

  Russia

A pliosaurid. The type species is Makhaira rossica.

Vegasaurus[16]

Gen. et sp. nov

Valid

O’Gorman et al.

Late Cretaceous (early Maastrichtian)

Snow Hill Island Formation

Antarctica (Vega Island)

An elasmosaurid plesiosaur. The type species is Vegasaurus molyi.

Wangosaurus[17]

Gen. et sp. nov

Valid

Ma et al.

Middle Triassic (Ladinian)

Falang Formation

  China

A pistosauroid. The type species is Wangosaurus brevirostris.

Lepidosaurs edit

Research edit

  • A phylogenetic analysis of living and fossil squamate relationships, based on morphological and molecular data, is published by Reeder et al. (2015).[18]
  • Miocene anoles from the Dominican amber, showing the habitat specializations also present in the extant Caribbean anoles, are described by Sherratt et al. (2015).[19]

New taxa edit

Rhynchocephalians edit

Name Novelty Status Authors Age Unit Location Notes Images

Clevosaurus sectumsemper[20]

Sp. nov

Valid

Klein et al.

Late Triassic (Rhaetian)

  United Kingdom

A clevosaurid sphenodontian, a species of Clevosaurus.

Lizards edit

Name Novelty Status Authors Age Unit Location Notes Images

Archaerhineura[21]

Gen. et sp. nov

Valid

Longrich et al.

Late Palaeocene

Polecat Bench Formation

  United States

A rhineurid amphisbaenian. The type species is Archaerhineura mephitis.

Asagaolacerta[22]

Gen. et sp. nov

Valid

Evans & Matsumoto

Early Cretaceous

Kuwajima Formation

  Japan

A lizard of uncertain phylogenetic placement. The type species is Asagaolacerta tricuspidens.

Asprosaurus[23]

Gen. et sp. nov

Valid

Park, Evans & Huh

Late Cretaceous

  South Korea

A member of Anguimorpha, probably a member of Monstersauria. The type species is Asprosaurus bibongriensis.

Babibasiliscus[24]

Gen. et sp. nov

Valid

Conrad

Eocene (approximately 48 Ma)

Bridger Formation

  United States

A member of Corytophanidae. The type species is Babibasiliscus alxi.

Barbatteius[25]

Gen. et sp. nov.

Valid

Venczel & Codrea

Late Cretaceous (early Maastrichtian)

Haţeg Basin

  Romania

Originally assigned to the family Teiidae, but subsequently transferred to the separate family Barbatteiidae within the group Teiioidea.[26] The type species is Barbatteius vremiri.

Cadurcogekko verus[27]

Sp. nov

Valid

Bolet et al.

Eocene

  France

A gekkotan lizard, a species of Cadurcogekko.

Chromatogenys[28]

Gen. et sp. nov

Valid

Makádi & Nydam

Late Cretaceous (Santonian)

  Hungary

A member of Scincomorpha of uncertain phylogenetic placement. The type species is Chromatogenys tiliquoides.

Chthonophis[21]

Gen. et sp. nov

Valid

Longrich et al.

Probably early Palaeocene

Bug Creek Anthills, Fort Union Formation

  United States

An amphisbaenian, the only member of the new family Chthonophidae. The type species is Chthonophis subterraneus.

Cuvieribaena[29]

Gen. et sp. nov

Valid

Čerňanský, Augéc & Rage

Eocene (Bartonian)

  France

A blanid amphisbaenian. The type species is Cuvieribaena carlgansi.

Dryadissector[30]

Gen. et sp. nov

Valid

Wick, Lehman & Brink

Late Cretaceous (early Campanian)

Aguja Formation

  United States

A member of Varanoidea. The type species is Dryadissector shilleri.

Gekkomimus[27]

Gen. et comb. nov

Valid

Bolet et al.

Eocene

  France

A skink; a new genus for "Cadurcogekko" rugosus Augé (2005).

Gueragama[31]

Gen. et sp. nov

Valid

Simões et al.

Late Cretaceous (Turonian-Campanian)

Goio-Erê Formation

  Brazil

The first member of Acrodonta (a relative of agamids and chameleons) described from South America. The type species is Gueragama sulamericana.

Hakuseps[22]

Gen. et sp. nov

Valid

Evans & Matsumoto

Early Cretaceous

  Japan

A squamate of uncertain phylogenetic placement. The type species is Hakuseps imberis.

Kuroyuriella[22]

Gen. et sp. nov

Valid

Evans & Matsumoto

Early Cretaceous

Kuwajima Formation

  Japan

A lizard of uncertain phylogenetic placement. The type species is Kuroyuriella mikikoi.

Norellius[32]

Gen. et sp. nov

Valid

Conrad & Daza

Early Cretaceous (probably approximately 130 Mya)

Öösh Basin

  Mongolia

A member of Gekkonomorpha of uncertain phylogenetic placement. The type species is Norellius nyctisaurops.

Ophisaurus holeci[33]

Sp. nov

Valid

Klembara

Miocene

  Czech Republic
  Germany[34][35]

A glass lizard.

Pachygenys adachii[36]

Sp. nov

Valid

Ikeda, Ota & Saegusa

Early Cretaceous

Sasayama Group

  Japan

A squamate of uncertain phylogenetic placement, a species of Pachygenys.

Phosphorosaurus ponpetelegans[37]

Sp. nov

Valid

Konishi et al.

Late Cretaceous (Maastrichtian)

  Japan

A halisaurine mosasaur, a species of Phosphorosaurus.

Plioplatecarpus peckensis[38]

Sp. nov

Valid

Cuthbertson & Holmes

Late Cretaceous (Campanian)

Bearpaw Formation

  United States

A mosasaur, a species of Plioplatecarpus.

Pseudopus rugosus[33]

Sp. nov

Valid

Klembara

Early Miocene

  Czech Republic

A relative of the sheltopusik.

Tetrapodophis[39]

Gen. et sp. nov

Valid

Martill, Tischlinger & Longrich

Early Cretaceous (Aptian)

Crato Formation

  Brazil

A squamate reptile of uncertain phylogenetic placement; originally classified as an early, four-legged snake,[39] but subsequently argued to be a dolichosaurid.[40][41] The type species is Tetrapodophis amplectus.

Snakes edit

Name Novelty Status Authors Age Unit Location Notes Images

Diablophis[42]

Gen. et comb. nov

Valid

Caldwell et al.

Late Jurassic

Morrison Formation

  United States

A basal snake; a new genus for "Parviraptor" gilmorei Evans (1996).

Eophis[42]

Gen. et sp. nov

Valid

Caldwell et al.

Middle Jurassic (Bathonian)

Forest Marble

  United Kingdom

A basal snake. The type species is Eophis underwoodi.

Portugalophis[42]

Gen. et sp. nov

Valid

Caldwell et al.

Late Jurassic (Kimmeridgian)

Camadas de Guimarota

  Portugal

A basal snake. The type species is Portugalophis lignites.

Renenutet[43]

Gen. et sp. nov

Valid

Mccartney & Seiffert

Eocene (Priabonian)

  Egypt

A member of Colubroidea. The type species is Renenutet enmerwer.

Turtles edit

Name Novelty Status Authors Age Unit Location Notes Images

Adocus sengokuensis[44]

Sp. nov

Valid

Sonoda et al.

Early Cretaceous

Sengoku Formation

  Japan

A species of Adocus.

Arvinachelys[45]

Gen. et sp. nov

Valid

Lively

Late Cretaceous (Campanian)

Kaiparowits Formation

  United States

A member of Baenidae. The type species is Arvinachelys goldeni.

Bairdemys thalassica[46]

Sp. nov

Valid

Ferreira et al.

Miocene

Capadare Formation

  Venezuela

A podocnemidid belonging to the clade Stereogenyina, a species of Bairdemys.

Baltemys velogastros[47]

Sp. nov

Valid

Lichtig & Lucas

Eocene (early Bridgerian)

Huerfano Formation

  United States

A member of Kinosternidae, a species of Baltemys.

Corsochelys bentleyi[48]

Sp. nov

Valid

Schwimmer et al.

Late Cretaceous

  United States

A member of Dermochelyidae, a species of Corsochelys.

Desmatochelys padillai[49]

Sp. nov

Valid

Cadena & Parham

Early Cretaceous (late Barremian-early Aptian)

Paja Formation

  Colombia

A member of Protostegidae, species of Desmatochelys.

 
Desmatochelys padillai

Gaffneylania[50]

Gen. et sp. nov

Valid

Sterli, de la Fuente & Krause

Middle Eocene

Sarmiento Formation

  Argentina

A member of Meiolaniidae. The type species is Gaffneylania auricularis.

Gobiapalone palaeocenica[51]

Sp. nov

Valid

Danilov et al.

Paleocene

Bugin Tsav Basin

  Mongolia

A member of Trionychidae, a species of Gobiapalone. The genus Gobiapalone was considered to be a junior synonym of the genus Kuhnemys by Georgalis & Joyce (2017), though the authors maintained G. palaeocenica as a distinct species within the latter genus.[52]

Gomphochelys[53]

Gen. et sp. nov

Valid

Bourque et al.

Early Wasatchian

Willwood Formation

  United States

A member of Dermatemydidae. The type species is Gomphochelys nanus.

Judithemys russelli[54]

Sp. nov

Valid

Brinkman

Late Cretaceous

  Canada

A member of (likely non-monophyletic) Macrobaenidae, a species of Judithemys.

Jurassichelon[55]

Gen. et sp. nov

Valid

Pérez-García

Late Jurassic (early Tithonian)

  France

A basal member of Eucryptodira. The type species is Jurassichelon oleronensis.

Khunnuchelys lophorhothon[56]

Sp. nov

Valid

Danilov et al.

Late Cretaceous (Santonian or early Campanian)

Bostobe Formation

  Kazakhstan

A trionychine trionychid, a species of Khunnuchelys.

Kimurachelys[57]

Gen. et sp. nov

Valid

Brinkman et al.

Late Cretaceous (late Campanian)

Dinosaur Park Formation

  Canada

A chelonioid turtle. The type species is Kimurachelys slobodae.

Leyvachelys[58]

Gen. et sp. nov

Valid

Cadena

Early Cretaceous (Barremian-Aptian)

Glen Rose Formation
Paja Formation

  Colombia
  United States

A member of Pan-Cryptodira belonging to the family Sandownidae. The type species is Leyvachelys cipadi.

Mauremys oshiroi[59]

Sp. nov.

Valid

Takahashi et al.

Late Pleistocene

Tomori Amaga Cave

  Japan

A pond turtle.

Neurankylus hutchisoni[60]

Sp. nov

Valid[61]

Lively

Late Cretaceous (Campanian)

Kaiparowits Formation

  United States

A member of Baenidae, a species of Neurankylus.

Neurankylus utahensis[60]

Sp. nov

Valid[61]

Lively

Late Cretaceous (Campanian)

Kaiparowits Formation

  United States

A member of Baenidae, a species of Neurankylus.

Perochelys[62]

Gen. et sp. nov.

Valid

Li, Joyce & Liu

Early Cretaceous (Aptian)

Jiufotang Formation

  China

A soft-shelled turtle of uncertain phylogenetic placement; it might be a stem- or a crown-trionychid. The type species is Perochelys lamadongensis.

Phunoichelys[63]

Gen. et sp. nov.

Valid

Tong et al.

Late Jurassic

Phu Kradung Formation

  Thailand

A xinjiangchelyid eucryptodiran. The type species is Phunoichelys thirakhupti.

Portlandemys gracilis[64]

Sp. nov.

Valid

Anquetin, Püntener & Billon-Bruyat

Late Jurassic (late Kimmeridgian)

Reuchenette Formation

   Switzerland

A plesiochelyid eucryptodiran, a species of Portlandemys.

Sternotherus bonevalleyensis[65]

Sp. nov.

Valid

Bourque & Schubert

Late Hemphillian

  United States

A musk turtle.

Sternotherus palaeodorus[65]

Sp. nov.

Valid

Bourque & Schubert

Late Hemphillian

  United States

A musk turtle.

Taraschelon[66]

Gen. et comb. nov

Valid[67]

Pérez-García

Early Oligocene

  France

A tortoise; a new genus for "Testudo" gigas Bravard (1844).

Testudo brevitesta[68]

Sp. nov

Valid[69]

Vlachos & Tsoukala

Late Pliocene (early Villafranchian)

  Greece

A tortoise, a species of Testudo.

Thalassemys bruntrutana[70]

Sp. nov

Valid

Püntener, Anquetin & Billon-Bruyat

Late Jurassic (late Kimmeridgian)

Reuchenette Formation

   Switzerland

A basal member of Pancryptodira, a species of Thalassemys.

Toremys[71]

Gen. et sp. nov.

Valid

Pérez-García et al.

Early Cretaceous (early Albian)

  Spain

A pleurosternid paracryptodiran. The type species is Toremys cassiopeia.

"Trionyx" jixiensis[72]

Sp. nov.

Valid

Li et al.

Early Cretaceous

Chengzihe Formation

  China

A member of Trionychinae of uncertain phylogenetic placement.

Xiaochelys[73]

Gen. et sp. nov

Valid

Zhou & Rabi

Early Cretaceous (Barremian-early Aptian)

Yixian Formation

  China

A member of Sinemydidae. The type species is Xiaochelys ningchengensis.

Archosauriformes edit

Archosaurs edit

Other edit

Other reptiles edit

Name Novelty Status Authors Age Unit Location Notes Images

Eohyosaurus[75]

Gen. et sp. nov

Valid

Butler et al.

Middle Triassic (early Anisian)

Burgersdorp Formation

  South Africa

A basal rhynchosaur. The type species is Eohyosaurus wolvaardti.

Erpetonyx[76]

Gen. et sp. nov

Valid

Modesto et al.

Carboniferous (Gzhelian)

Egmont Bay Formation

  Canada

A relative of bolosaurids. The type species is Erpetonyx arsenaultorum.

Glaurung[77]

Gen. et sp. nov

Valid

Bulanov & Sennikov

Late Permian

  Germany

A member of Weigeltisauridae. The type species is Glaurung schneideri.

Opisthodontosaurus[78]

Gen. et sp. nov

Valid

Reisz et al.

Early Permian

Garber Formation

  United States

A member of Captorhinidae. The type species is Opisthodontosaurus carrolli.

Pappochelys[79]

Gen. et sp. nov

Valid

Schoch & Sues

Middle Triassic (Ladinian)

Erfurt Formation

  Germany

A stem-turtle. The type species is Pappochelys rosinae.

 
Pappochelys

"Thalassodromeus" sebesensis[80]

Sp. nov

Valid

Grellet-Tinner & Codrea

Late Cretaceous (Maastrichtian)

  Romania

A reptile of uncertain phylogenetic placement; initially classified as a pterosaur and a species of Thalassodromeus,[80] but subsequently argued to be a turtle and a junior synonym of Kallokibotion bajazidi.[81]

References edit

  1. ^ Cheng Ji; Da-Yong Jiang; Ryosuke Motani; Olivier Rieppel; Wei-Cheng Hao & Zuo-Yu Sun (2015). "Phylogeny of the Ichthyopterygia incorporating recent discoveries from South China". Journal of Vertebrate Paleontology. 36 (1): e1025956. doi:10.1080/02724634.2015.1025956. S2CID 85621052.
  2. ^ Jeremy E. Martin; Peggy Vincent; Guillaume Suan; Tom Sharpe; Peter Hodges; Matt Williams; Cindy Howells & Valentin Fischer (2015). "A mysterious giant ichthyosaur from the lowermost Jurassic of Wales". Acta Palaeontologica Polonica. 60 (4): 837–842. doi:10.4202/app.00062.2014.
  3. ^ Ryosuke Motani; Da-Yong Jiang; Guan-Bao Chen; Andrea Tintori; Olivier Rieppel; Cheng Ji & Jian-Dong Huang (2015). "A basal ichthyosauriform with a short snout from the Lower Triassic of China". Nature. 517 (7535): 485–488. Bibcode:2015Natur.517..485M. doi:10.1038/nature13866. PMID 25383536. S2CID 4392798.
  4. ^ Stephen L. Brusatte; Mark T. Young; Thomas J. Challands; Neil D. L. Clark; Valentin Fischer; Nicholas C. Fraser; Jeff J. Liston; Colin C. J. MacFadyen; Dugald A. Ross; Stig Walsh; Mark Wilkinson (2015). "Ichthyosaurs from the Jurassic of Skye, Scotland" (PDF). Scottish Journal of Geology. 51 (1): 43–55. doi:10.1144/sjg2014-018. hdl:2268/176434. S2CID 54614526.
  5. ^ Xiao-hong Chen; Ryosuke Motani; Long Cheng; Da-yong Jiang & Olivier Rieppel (2015). "A New Specimen of Carroll's Mystery Hupehsuchian from the Lower Triassic of China". PLOS ONE. 10 (5): e0126024. Bibcode:2015PLoSO..1026024C. doi:10.1371/journal.pone.0126024. PMC 4446317. PMID 26017585.
  6. ^ Dean R. Lomax; Judy A. Massare (2015). "A new species of Ichthyosaurus from the Lower Jurassic of West Dorset, England, U.K." Journal of Vertebrate Paleontology. 35 (2): e903260. doi:10.1080/02724634.2014.903260. S2CID 85745787.
  7. ^ Erin E. Maxwell; Daniel Dick; Santiago Padilla; Mary Luz Parra (2015). "A new ophthalmosaurid ichthyosaur from the Early Cretaceous of Colombia". Papers in Palaeontology. 2 (1): 59–70. doi:10.1002/spp2.1030. S2CID 128464793.
  8. ^ Nicole Klein; James M. Neenan; Torsten M. Scheyer; Eva Maria Griebeler (2015). "Growth patterns and life-history strategies in Placodontia (Diapsida: Sauropterygia)". Royal Society Open Science. 2 (7): 140440. Bibcode:2015RSOS....240440K. doi:10.1098/rsos.140440. PMC 4632572. PMID 26587259.
  9. ^ Ryosuke Motani; Da-yong Jiang; Olivier Rieppel; Yi-fan Xue; Andrea Tintori (2015). "Adult sex ratio, sexual dimorphism and sexual selection in a Mesozoic reptile". Proceedings of the Royal Society B: Biological Sciences. 282 (1815): 20151658. doi:10.1098/rspb.2015.1658. PMC 4614760. PMID 26378218.
  10. ^ Judyth Sassoon; Davide Foffa; Ryan Marek (2015). "Dental ontogeny and replacement in Pliosauridae". Royal Society Open Science. 2 (11): 150384. Bibcode:2015RSOS....250384S. doi:10.1098/rsos.150384. PMC 4680613. PMID 26715998.
  11. ^ Andrea Cau; Federico Fanti (2015). "High evolutionary rates and the origin of the Rosso Ammonitico Veronese Formation (Middle-Upper Jurassic of Italy) reptiles". Historical Biology: An International Journal of Paleobiology. 28 (7): 952–962. doi:10.1080/08912963.2015.1073726. S2CID 86528030.
  12. ^ Adam S. Smith (2015). "Reassessment of 'Plesiosaurus' megacephalus (Sauropterygia: Plesiosauria) from the Triassic-Jurassic boundary, UK". Palaeontologia Electronica. 18 (1): Article number 18.1.20A.
  13. ^ R. Araújo; M.J. Polcyn; A.S. Schulp; O. Mateus; L.L. Jacobs; A. Olímpio Gonçalves; M.-L. Morais (2015). "A new elasmosaurid from the early Maastrichtian of Angola and the implications of girdle morphology on swimming style in plesiosaurs". Netherlands Journal of Geosciences. 94 (1): 109–120. doi:10.1017/njg.2014.44. S2CID 86616531.
  14. ^ Qing-Hua Shang; Chun Li (2015). "A new small-sized eosauropterygian (Diapsida: Sauropterygia) from the Middle Triassic of Luoping, Yunnan, southwestern China". Vertebrata PalAsiatica. 53 (4): 265–280. doi:10.19615/j.cnki.1000-3118.2015.04.001.
  15. ^ Valentin Fischer; Maxim S. Arkhangelsky; Ilya M. Stenshin; Gleb N. Uspensky; Nikolay G. Zverkov; Roger B. J. Benson (2015). "Peculiar macrophagous adaptations in a new Cretaceous pliosaurid". Royal Society Open Science. 2 (12): 150552. Bibcode:2015RSOS....250552F. doi:10.1098/rsos.150552. PMC 4807462. PMID 27019740.
  16. ^ José P. O’Gorman; Leonardo Salgado; Eduardo B. Olivero; Sergio A. Marenssi (2015). "Vegasaurus molyi, gen. et sp. nov. (Plesiosauria, Elasmosauridae), from the Cape Lamb Member (lower Maastrichtian) of the Snow Hill Island Formation, Vega Island, Antarctica, and remarks on Wedellian Elasmosauridae". Journal of Vertebrate Paleontology. 35 (3): e931285. doi:10.1080/02724634.2014.931285. hdl:11336/53416. S2CID 128965534.
  17. ^ Le-Tian Ma; Da-Yong Jiang; Olivier Rieppel; Ryosuke Motani; Andrea Tintori (2015). "A new pistosauroid (Reptilia, Sauropterygia) from the late Ladinian Xingyi marine reptile level, southwestern China". Journal of Vertebrate Paleontology. 35 (1): e881832. doi:10.1080/02724634.2014.881832. S2CID 130696958.
  18. ^ Tod W. Reeder; Ted M. Townsend; Daniel G. Mulcahy; Brice P. Noonan; Perry L. Wood Jr.; Jack W. Sites Jr.; John J. Wiens (2015). "Integrated Analyses Resolve Conflicts over Squamate Reptile Phylogeny and Reveal Unexpected Placements for Fossil Taxa". PLOS ONE. 10 (3): e0118199. Bibcode:2015PLoSO..1018199R. doi:10.1371/journal.pone.0118199. PMC 4372529. PMID 25803280.
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