2014 in reptile paleontology

This list of fossil reptiles described in 2014 is a list of new taxa of fossil reptiles that were described during the year 2014, as well as other significant discoveries and events related to reptile paleontology that occurred in 2014.

List of years in reptile paleontology
In science
2011
2012
2013
2014
2015
2016
2017
In paleobotany
2011
2012
2013
2014
2015
2016
2017
In paleontology
2011
2012
2013
2014
2015
2016
2017
In arthropod paleontology
2011
2012
2013
2014
2015
2016
2017
In paleoentomology
2011
2012
2013
2014
2015
2016
2017
In paleomalacology
2011
2012
2013
2014
2015
2016
2017
In archosaur paleontology
2011
2012
2013
2014
2015
2016
2017
In mammal paleontology
2011
2012
2013
2014
2015
2016
2017
In paleoichthyology
2011
2012
2013
2014
2015
2016
2017

Ichthyopterygians edit

Research edit

New taxa edit

Name Novelty Status Authors Age Unit Location Notes Images

Janusaurus[2]

Gen. et sp. nov

Valid

Roberts et al.

Late Jurassic

Agardhfjellet Formation

  Norway

An ophthalmosaurid ichthyosaur. The type species is Janusaurus lundi. Zverkov & Prilepskaya (2019) considered Janusaurus to be a junior synonym of the genus Arthropterygius, though the authors maintained J. lundi as a distinct species within the latter genus;[3] Delsett et al. (2019) rejected this synonymy.[4]

Leninia[5]

Gen. et sp. nov

Valid

Fischer et al.

Early Cretaceous (early Aptian)

  Russia

An ophthalmosaurine ophthalmosaurid ichthyosaur. The type species is Leninia stellans.

Sisteronia[6]

Gen. et sp. nov

Valid

Fischer et al.

Cretaceous (early Albian to early Cenomanian)

Cambridge Greensand
Gault Formation
Marnes Bleues Formation

  France
  United Kingdom

A platypterygiine ophthalmosaurid ichthyosaur. The type species is Sisteronia seeleyi.

Undorosaurus trautscholdi[7]

Sp. nov

Valid

Arkhangelsky & Zverkov

Late Jurassic

  Russia

An ophthalmosaurine ophthalmosaurid ichthyosaur, a species of Undorosaurus.

Sauropterygians edit

Name Novelty Status Authors Age Unit Location Notes Images

Aristonectes quiriquinensis[8]

Sp. nov

Valid

Otero et al.

Late Cretaceous (late Maastrichtian)

Quiriquina Formation

  Chile

A plesiosaur, a species of Aristonectes.

Cymatosaurus erikae[9]

Sp. nov

Valid

Maisch

Middle Triassic (earliest Anisian)

Röt Formation

  Germany

A species of Cymatosaurus.

Majiashanosaurus[10]

Gen. et sp. nov

Valid

Jiang et al.

Early Triassic (Olenekian)

Nanlinghu Formation

  China

A basal member of Eosauropterygia. The type species is Majiashanosaurus discocoracoidis.

Nothosaurus zhangi[11]

Sp. nov

Valid

Liu et al.

Middle Triassic (Anisian)

Guanling Formation

  China

A nothosaur, a species of Nothosaurus.

Odoiporosaurus[12]

Gen. et sp. nov

Valid

Renesto, Binelli & Hagdorn

Middle Triassic (Anisian)

Besano Formation

  Italy

A pachypleurosaur. The type species is Odoiporosaurus teruzzii.

Pararcus[13]

Gen. et sp. nov

Valid

Klein & Scheyer

Middle Triassic (early Anisian)

Vossenveld Formation

  Netherlands

A placodont. The type species is Pararcus diepenbroeki.

Pliosaurus patagonicus[14]

Sp. nov

Valid

Gasparini & O'Gorman

Late Jurassic (middle Tithonian)

Vaca Muerta Formation

  Argentina

A species of Pliosaurus.

Lepidosaurs edit

Newly named rhynchocephalians edit

Name Novelty Status Authors Age Unit Location Notes Images

Kawasphenodon peligrensis[15]

Sp. nov

Valid

Apesteguia, Gómez & Rougier

Palaeocene

Salamanca Formation

  Argentina

An opisthodontian sphenodontid, a species of Kawasphenodon.

Priosphenodon minimus[16]

Sp. nov

Valid

Apesteguia & Carballido

Early Cretaceous

Cerro Barcino Formation

  Argentina

An eilenodontine sphenodontid, a species of Priosphenodon.

Newly named lizards edit

Name Novelty Status Authors Age Unit Location Notes Images

Blanus mendezi[17]

Sp. nov

Valid

Bolet et al.

Miocene (11.6 Mya)

Vallès-Penedès Basin

  Spain

An amphisbaenian, a species of Blanus.

Blanus thomaskelleri[18]

Sp. nov

Valid[19]

Čerňanský, Rage & Klembara

Early Miocene

  Germany

An amphisbaenian, a species of Blanus.

Calanguban[20]

Gen. et sp. nov

Valid[21]

Simões, Caldwell & Kellner

Early Cretaceous

Crato Formation

  Brazil

A scleroglossan lizard with scincomorph affinities. The type species is Calanguban alamoi.

Dakotaseps[22]

Nom. nov

Valid

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

  United States

A member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade;[23] a replacement name for Dakotasaurus Nydam (2013) (preoccupied).

Funiusaurus[24]

Gen. et sp. nov

Valid

Xu et al.

Late Cretaceous

Qiupa Formation

  China

A member of Polyglyphanodontidae. The type species is Funiusaurus luanchuanensis.

Maioricalacerta[25]

Gen. et sp. nov

Valid

Bailon et al.

Early Pliocene (Zanclean)

  Spain

A member of Lacertidae. The type species is Maioricalacerta rafelinensis.

Pyrenasaurus[26]

Gen. et sp. nov

Valid

Bolet & Augé

Late Eocene

  France
  Spain

A member of Scincoidea, possibly a skink. The type species is Pyrenasaurus evansae.

Newly named snakes edit

Name Novelty Status Authors Age Unit Location Notes Images

Adinophis[27]

Gen. et sp. nov

Valid

Pritchard et al.

Late Cretaceous (Maastrichtian)

Maevarano Formation

  Madagascar

A member of Madtsoiidae. The type species is Adinophis fisaka.

Indophis fanambinana[27]

Sp. nov

Valid

Pritchard et al.

Late Cretaceous (Maastrichtian)

Maevarano Formation

  Madagascar

A member of Nigerophiidae, a species of Indophis.

Rukwanyoka[28]

Gen. et sp. nov

Valid

McCartney et al.

Oligocene

Nsungwe Formation

  Tanzania

The most ancient booid snake and boa from Africa. The type species is Rukwanyoka holmani.

Seismophis[29]

Gen. et sp. nov

Valid

Hsiou et al.

Late Cretaceous (Cenomanian)

Alcântara Formation

  Brazil

A snake of uncertain phylogenetic placement, possibly a relative of Najash rionegrina. The type species is Seismophis septentrionalis.

Turtles edit

Research edit

Newly named turtles edit

Name Novelty Status Authors Age Unit Location Notes Images

Alienochelys[31]

Gen. et sp. nov

Valid

de Lapparent de Broin et al.

Late Cretaceous (Maastrichtian)

Oulad Abdoun Basin

  Morocco

A sea turtle, a member of Dermochelyoidae. The type species is Alienochelys selloumi.

Allaeochelys libyca[32]

Sp. nov

Valid

Havlik, Joyce & Böhme

Miocene (Langhian)

  Libya

A relative of the pig-nosed turtle, a species of the (possibly paraphyletic) genus Allaeochelys.

Ashleychelys[33]

Gen. et sp. nov

Valid

Weems & Sanders

Oligocene

  United States

A pancheloniid sea turtle. The type species is Ashleychelys palmeri.

Atolchelys[34]

Gen. et sp. nov

Valid

Romano et al.

Early Cretaceous (Barremian)

Morro do Chaves Formation

  Brazil

A bothremydid pleurodiran. The type species is Atolchelys lepida.

Brodiechelys royoi[35]

Sp. nov

Valid

Pérez-García, Gasulla & Ortega

Early Aptian

Arcillas de Morella Formation

  Spain

A xinjiangchelyid, a species of Brodiechelys.

Cheirogaster bacharidisi[36]

Sp. nov

Valid

Vlachos, Tsoukala & Corsini

Pliocene

Gonia Formation

  Greece

A tortoise, originally described as a species belonging to the genus Cheirogaster. Subsequently, transferred by Pérez-García & Vlachos (2014) to the genus Titanochelon.[37]

Eodortoka[38]

Gen. et sp. nov

Valid

Pérez-García, Gasulla & Ortega

Early Cretaceous (Aptian)

Arcillas de Morella Formation

  Spain

A dortokid, a member of the clade Pan-Pleurodira (containing living pleurodirans and all turtles that are more closely related to them than to cryptodirans). The type species is Eodortoka morellana.

Gobiapalone[39]

Gen. et comb. et sp. nov

Valid[40]

Danilov et al.

Late Cretaceous (Cenomanian to Maastrichtian)

Barun Goyot Formation
Bayan Shireh Formation
Nemegt Formation

  Mongolia

A trionychine trionychid. The type species is "Amyda" orlovi Khosatzky (1976); genus also contains new species Gobiapalone breviplastra. The genus Gobiapalone was considered to be a junior synonym of the genus Kuhnemys by Georgalis & Joyce (2017), though the authors maintained G. orlovi and G. breviplastra as distinct species within the latter genus.[41]

Hylaeochelys kappa[42]

Sp. nov

Valid

Pérez-García & Ortega

Late Jurassic (Tithonian)

Freixial Formation

  Portugal

A basal member of Eucryptodira, a species of Hylaeochelys.

Judithemys kranzi[43]

Sp. nov

Valid

Weems

Paleocene (early Thanetian)

Aquia Formation

  United States

A member of (likely paraphyletic) group "Macrobaenidae", a species of Judithemys.

Nemegtemys[39]

Gen. et sp. nov

Valid[40]

Danilov et al.

Late Cretaceous (Maastrichtian)

Nemegt Formation

  Mongolia

A cyclanorbine trionychid. The type species is Nemegtemys conflata.

Osonachelus[44]

Gen. et sp. nov

Valid

de Lapparent de Broin et al.

Eocene (late Bartonian)

Vic-Manlleu Marls Formation

  Spain

A cheloniid sea turtle. The type species is Osonachelus decorata.

Procolpochelys charlestonensis[33]

Sp. nov

Valid

Weems & Sanders

Oligocene

  United States

A pancheloniid sea turtle; a species of Procolpochelys.

Riodevemys[45]

Gen. et sp. nov

Valid[46]

Pérez-García, Royo-Torres & Cobos

Late Jurassic

  Spain

A pleurosternid paracryptodiran. The type species is Riodevemys inumbragigas.

Tacuarembemys[47]

Gen. et sp. nov

Valid

Perea et al.

Late Jurassic or earliest Cretaceous

Tacuarembó Formation

  Uruguay

A turtle of uncertain phylogenetic placement. The type species is Tacuarembemys kusterae.

Titanochelon[37]

Gen. et comb. nov

Valid

Pérez-García & Vlachos

Early Miocene to early Pleistocene

  Austria
  Bulgaria
  France
  Germany
  Greece
  Portugal
  Spain
   Switzerland
  Turkey
  Malta?

A tortoise; a new genus for "Testudo" bolivari Hernández-Pacheco (1917). Genus also contains "Testudo" eurysternum Gervais (1848–1852), "Testudo" ginsburgi de Broin (1977), "Testudo" vitodurana Biedermann (1862), "Cheirogaster" steinbacheri Karl (1996), "Testudo" leberonensis Depéret (1890), "Testudo" schafferi Szalai (1931), "Testudo" perpiniana Depéret (1885) and "Cheirogaster" bacharidisi Vlachos et al. (2014). Genus might also contain "Testudo" gymnesicus Bate (1914).

‘Trionyx’ baynshirensis[39]

Sp. nov

Valid[40]

Danilov et al.

Late Cretaceous (Cenomanian to Santonian)

Bayan Shireh Formation

  Mongolia

A trionychine trionychid, a species of Trionyx sensu lato.

‘Trionyx’ dissolutus[48]

Sp. nov

Valid

Vitek & Danilov

Late Cretaceous (Cenomanian)

  Uzbekistan

A trionychid, a species of Trionyx sensu lato.

‘Trionyx’ gilbentuensis[39]

Sp. nov

Valid[40]

Danilov et al.

Late Cretaceous (Maastrichtian)

Nemegt Formation

  Mongolia

A trionychine trionychid, a species of Trionyx sensu lato.

‘Trionyx’ gobiensis[39]

Sp. nov

Valid[40]

Danilov et al.

Late Cretaceous (Maastrichtian)

Nemegt Formation

  Mongolia

A trionychine trionychid, a species of Trionyx sensu lato.

‘Trionyx’ shiluutulensis[39]

Sp. nov

Valid[40]

Danilov et al.

Late Cretaceous (Campanian)

  Mongolia

A trionychine trionychid, a species of Trionyx sensu lato.

Archosauromorphs edit

Archosaurs edit

Basal archosauromorphs edit

Research edit

  • A redescription of Tasmaniosaurus triassicus is published by Ezcurra (2014).[49]
  • A revision of anatomy and phylogenetic relationships of the archosauriform Dorosuchus neoetus is published by Sookias et al. (2014).[50]
  • A revision of putative euparkeriids from the Triassic of China is published by Sookias et al. (2014).[51]

New taxa edit

Name Novelty Status Authors Age Unit Location Notes Images

Aenigmastropheus[52]

Gen. et sp. nov

Valid

Ezcurra, Scheyer & Bulter

Late Permian (Wuchiapingian)

Usili Formation

  Tanzania

A non-archosauriform archosauromorph, probably a protorosaurian. The type species is Aenigmastropheus parringtoni.

Garjainia madiba[53]

Sp. nov

Valid

Gower et al.

Early Triassic (late Olenekian)

Burgersdorp Formation

  South Africa

An erythrosuchid archosauriform, a species of Garjainia.

Hyperodapedon tikiensis[54]

Sp. nov

Valid[55]

Mukherjee & Ray

Late Triassic

Tiki Formation

  India

A rhynchosaur, a species of Hyperodapedon.

Proterosuchus goweri[56]

Sp. nov

Valid[57]

Ezcurra & Butler

Early Triassic

  South Africa

A proterosuchid archosauriform, a species of Proterosuchus.

Pseudochampsa[58]

Gen. et comb. nov

Valid

Trotteyn & Ezcurra

Late Triassic (late Carnian or earliest Norian)

Ischigualasto Formation

  Argentina

A proterochampsid archosauriform; a new genus for "Chanaresuchus" ischigualastensis Trotteyn, Martínez & Alcober (2012).

Other reptiles edit

Name Novelty Status Authors Age Unit Location Notes Images

Abyssomedon[59]

Gen. et sp. nov

Valid

Macdougall & Reisz

Early Permian

Garber Formation

  United States

A nyctiphruretid parareptile. The type species is Abyssomedon williamsi.

Atopodentatus[60]

Gen. et sp. nov

Valid

Cheng et al.

Middle Triassic (Anisian)

Guanling Formation

  China

A marine reptile, probably a relative of sauropterygians. The type species is Atopodentatus unicus.

Delorhynchus cifellii[61]

Sp. nov

Valid

Reisz, Macdougall & Modesto

Early Permian

Garber Formation

  United States

A parareptile relative of lanthanosuchoids, a species of Delorhynchus.

Eohupehsuchus[62]

Gen. et sp. nov

Valid

Chen et al.

Early Triassic

Jialingjiang Formation

  China

A hupehsuchian. The type species is Eohupehsuchus brevicollis.

Horaffia[63]

Gen. et sp. nov

Valid

Klein & Hagdorn

Middle Triassic (Ladinian)

Erfurt Formation

  Germany

A marine diapsid reptile of uncertain phylogenetic placement. The type species is Horaffia kugleri.

Largocephalosaurus qianensis[64]

Sp. nov

Valid

Li et al.

Triassic

Guanling Formation

  China

A member of Saurosphargidae, a species of Largocephalosaurus.

Parahupehsuchus[65]

Gen. et sp. nov

Valid

Chen et al.

Early Triassic

Jialingjiang Formation

  China

A hupehsuchian. The type species is Parahupehsuchus longus.

References edit

  1. ^ Valentin Fischer; Maxim S. Arkhangelsky; Darren Naish; Ilya M. Stenshin; Gleb N. Uspensky & Pascal Godefroit (2014). "Simbirskiasaurus and Pervushovisaurus reassessed: implications for the taxonomy and cranial osteology of Cretaceous platypterygiine ichthyosaurs" (PDF). Zoological Journal of the Linnean Society. 171 (4): 822–841. doi:10.1111/zoj.12158. S2CID 67765108.
  2. ^ Aubrey Jane Roberts; Patrick Scott Druckenmiller; Glenn-Peter Sætre & Jørn Harald Hurum (2014). "A New Upper Jurassic Ophthalmosaurid Ichthyosaur from the Slottsmøya Member, Agardhfjellet Formation of Central Spitsbergen". PLOS ONE. 9 (8): e103152. Bibcode:2014PLoSO...9j3152R. doi:10.1371/journal.pone.0103152. PMC 4118863. PMID 25084533.
  3. ^ Nikolay G. Zverkov; Natalya E. Prilepskaya (2019). "A prevalence of Arthropterygius (Ichthyosauria: Ophthalmosauridae) in the Late Jurassic—earliest Cretaceous of the Boreal Realm". PeerJ. 7: e6799. doi:10.7717/peerj.6799. PMC 6497043. PMID 31106052.
  4. ^ Lene L. Delsett; Aubrey J. Roberts; Patrick S. Druckenmiller; Jørn H. Hurum (2019). "Osteology and phylogeny of Late Jurassic ichthyosaurs from the Slottsmøya Member Lagerstätte (Spitsbergen, Svalbard)". Acta Palaeontologica Polonica. 64 (4): 717–743. doi:10.4202/app.00571.2018. hdl:10852/76089. S2CID 210294877.
  5. ^ Valentin Fischer; Maxim S. Arkhangelsky; Gleb N. Uspensky; Ilya M. Stenshin & Pascal Godefroit (2014). "A new Lower Cretaceous ichthyosaur from Russia reveals skull shape conservatism within Ophthalmosaurinae". Geological Magazine. 151 (1): 60–70. Bibcode:2014GeoM..151...60F. doi:10.1017/S0016756812000994. hdl:2268/137853. S2CID 49470362.
  6. ^ Valentin Fischer; Nathalie Bardet; Myette Guiomar & Pascal Godefroit (2014). "High Diversity in Cretaceous Ichthyosaurs from Europe Prior to Their Extinction". PLOS ONE. 9 (1): e84709. Bibcode:2014PLoSO...984709F. doi:10.1371/journal.pone.0084709. PMC 3897400. PMID 24465427.
  7. ^ M.S. Arkhangelsky & N.G. Zverkov (2014). "On a new ichthyosaur of the genus Undorosaurus" (PDF). Proceedings of the Zoological Institute of the Russian Academy of Sciences. 318 (3): 187–196.
  8. ^ Rodrigo A. Otero; Sergio Soto-Acuña; Frank Robin O'Keefe; José P. O’Gorman; Wolfgang Stinnesbeck; Mario E. Suárez; David Rubilar-Rogers; Christian Salazar; Luis Arturo Quinzio-Sinn (2014). "Aristonectes quiriquinensis, sp. nov., a new highly derived elasmosaurid from the upper Maastrichtian of central Chile". Journal of Vertebrate Paleontology. 34 (1): 100–125. doi:10.1080/02724634.2013.780953. hdl:11336/79718. S2CID 84729992.
  9. ^ Michael W. Maisch (2014). "A well preserved skull of Cymatosaurus (Reptilia: Sauropterygia) from the uppermost Buntsandstein (Middle Triassic) of Germany". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 272 (2): 213–224. doi:10.1127/0077-7749/2014/0407.
  10. ^ Da-Yong Jiang; Ryosuke Motani; Andrea Tintori; Olivier Rieppel; Guan-Bao Chen; Jian-Dong Huang; Rong Zhang; Zuo-Yu Sun; Cheng Ji (2014). "The Early Triassic eosauropterygian Majiashanosaurus discocoracoidis, gen. et sp. nov. (Reptilia, Sauropterygia), from Chaohu, Anhui Province, People's Republic of China". Journal of Vertebrate Paleontology. 34 (5): 1044–1052. doi:10.1080/02724634.2014.846264. S2CID 129828376.
  11. ^ Jun Liu; Shi-xue Hu; Olivier Rieppel; Da-yong Jiang; Michael J. Benton; Neil P. Kelley; Jonathan C. Aitchison; Chang-yong Zhou; Wen Wen; Jin-yuan Huang; Tao Xie; Tao Lv (2014). "A gigantic nothosaur (Reptilia: Sauropterygia) from the Middle Triassic of SW China and its implication for the Triassic biotic recovery". Scientific Reports. 4: Article number 7142. Bibcode:2014NatSR...4E7142L. doi:10.1038/srep07142. PMC 4245812. PMID 25429609.
  12. ^ Silvio Renesto; Giorgio Binelli; Hans Hagdorn (2014). "A new pachypleurosaur from the Middle Triassic Besano Formation of Northern Italy". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 271 (2): 151–168. doi:10.1127/0077-7749/2014/0382.
  13. ^ Nicole Klein & Torsten M. Scheyer (2014). "A new placodont sauropterygian from the Middle Triassic of the Netherlands". Acta Palaeontologica Polonica. 59 (4): 887–902. doi:10.4202/app.2012.0147. S2CID 54714765.
  14. ^ Zulma Gasparini; José P. O'Gorman (2014). "A new species of Pliosaurus (Sauropterygia, Plesiosauria) from the Upper Jurassic of northwestern Patagonia, Argentina". Ameghiniana. 51 (4): 269–283. doi:10.5710/amgh.03.04.2014.2225. hdl:11336/9372. S2CID 130194647.
  15. ^ Sebastian Apesteguia; Raúl O. Gómez; Guillermo W. Rougier (2014). "The youngest South American rhynchocephalian, a survivor of the K/Pg extinction". Proceedings of the Royal Society B: Biological Sciences. 281 (1792): 20140811. doi:10.1098/rspb.2014.0811. PMC 4150314. PMID 25143041.
  16. ^ Sebastian Apesteguia; Jose L. Carballido (2014). "A new eilenodontine (Lepidosauria, Sphenodontidae) from the Lower Cretaceous of central Patagonia". Journal of Vertebrate Paleontology. 34 (2): 303–317. doi:10.1080/02724634.2013.803974. hdl:11336/6524. S2CID 130381621.
  17. ^ Arnau Bolet; Massimo Delfino; Josep Fortuny; Sergio Almécija; Josep M. Robles; David M. Alba (2014). "An Amphisbaenian Skull from the European Miocene and the Evolution of Mediterranean Worm Lizards". PLOS ONE. 9 (6): e98082. Bibcode:2014PLoSO...998082B. doi:10.1371/journal.pone.0098082. PMC 4045672. PMID 24896828.
  18. ^ Andrej Čerňanský; Jean-Claude Rage; Jozef Klembara (2014). "The Early Miocene squamates of Amöneburg (Germany): the first stages of modern squamates in Europe". Journal of Systematic Palaeontology. 13 (2): 97–128. doi:10.1080/14772019.2014.897266. S2CID 140170242.
  19. ^ http://zoobank.org/References/426A24DC-719B-4D13-86BF-E2858F1F1D49 [dead link]
  20. ^ Tiago R. Simões; Michael W. Caldwell; Alexander W. A. Kellner (2014). "A new Early Cretaceous lizard species from Brazil, and the phylogenetic position of the oldest known South American squamates". Journal of Systematic Palaeontology. 13 (7): 601–614. doi:10.1080/14772019.2014.947342. S2CID 84446189.
  21. ^ "Archived copy". Archived from the original on 2019-05-10. Retrieved 2014-09-07.{{cite web}}: CS1 maint: archived copy as title (link)
  22. ^ Randall L. Nydam (2014). "Dakotaseps gen. nov., a replacement name for the lizard genus Dakotasaurus Nydam 2013, a junior homonym of the ichnotaxon Dakotasaurus Branson and Mehl 1932". Zootaxa. 3900 (1): 150. doi:10.11646/zootaxa.3900.1.11. PMID 25543731.
  23. ^ Randall L. Nydam (2013). "Lizards and Snakes from the Cenomanian through Campanian of Southern Utah: Filling the Gap in the Fossil Record of Squamata from the Late Cretaceous of the Western Interior of North America". In Alan L. Titus; Mark A. Loewen (eds.). At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah. Indiana University Press. pp. 370–423. ISBN 978-0-253-00896-1.
  24. ^ Li Xu; Xiaochun Wu; Junchang Lü; Songhai Jia; Jiming Zhang; Hanyong Pu; Xingliao Zhang (2014). "A New Lizard (Lepidosauria: Squamata) from the Upper Cretaceous of Henan, China". Acta Geologica Sinica - English Edition. 88 (4): 1041–1050. doi:10.1111/1755-6724.12271. S2CID 128552980.
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