The beetle family Scarabaeidae is made up of about 28 subfamilies containing more than 100 tribes. There are more than 2,500 genera and 35,000 described species in Scarabaeidae.[1]
The following subfamilies and tribes are in accordance with those in Catalog of Life (2023) and Dietz, Ahrens, et al. (2023). The tribe Hopliini is placed as a subfamily in Catalogue of Life, but is listed as a tribe of subfamily Melolonthinae here. The subfamilies Sericinae and Sericoidinae were suggested in Dietz, Ahrens, et al. (2023) and are included in the Catalogue of Life.[1][2] The subfamily Lichniinae is sometimes considered a tribe, Lichniini.[3]
Subfamily Aclopinae Blanchard, 1850 edit
- Aclopini Blanchard, 1850
- Holcorobeini Nikolajev, 1992
- Phaenognathini Iablokoff-Khnozorian, 1977
Subfamily Aegialiinae Castelnau, 1840 edit
- Aegialiini Castelnau, 1840
- Saprinianini Nikolajev, 2011
Subfamily Allidiostomatinae Arrow, 1940 edit
Subfamily Aphodiinae Leach, 1815 edit
- Aphodiini Leach, 1815
- Corythoderini Schmidt, 1910
- Eupariini Schmidt, 1910
- Hornietiini Minkina, 2020
- Odochilini Rakovič, 1987
- Odontolochini Stebnicka & Howden, 1996
- Psammodiini Mulsant, 1842
- Rhyparini Schmidt, 1910
- Stereomerini Howden & Storey, 1992
- Termitoderini Tangelder & Krikken, 1982
- Thinorycterini Semenov & Reichardt, 1925
- †Psammaegialiini Nikolajev, Wang & Zhang, 2014
Subfamily Aulonocneminae Janssens, 1946 edit
Subfamily Cetoniinae Leach, 1815 edit
- Cetoniini Leach, 1815
- Cremastocheilini Burmeister & Schaum, 1841
- Diplognathini Burmeister, 1842
- Goliathini Latreille, 1829
- Gymnetini Kirby, 1827
- Phaedimini Schoch, 1894
- Schizorhinini Burmeister, 1842
- Stenotarsiini Kraatz, 1880
- Taenioderini Mikšić, 1976
- Xiphoscelidini Krikken, 1984
Subfamily Chironinae Blanchard, 1845 edit
Subfamily Dynamopodinae Arrow, 1911 edit
Subfamily Dynastinae MacLeay, 1819 edit
- Agaocephalini Burmeister, 1847
- Cyclocephalini Castelnau, 1840
- Dynastini Macleay, 1819
- Hexodontini Lacordaire, 1855
- Oryctini Mulsant, 1842
- Oryctoderini Endrödi, 1966
- Pentodontini Mulsant, 1842
- Phileurini Burmeister, 1847
Subfamily Eremazinae Stebnicka, 1977 edit
- Eremazini Stebnicka, 1977
- †Yixianscarabaeini Nikolajev, 2015
Subfamily Lichniinae Burmeister, 1844 edit
Subfamily Melolonthinae MacLeay, 1819 edit
- Acomini Evans & Smith, 2020
- Ardellini Paulsen, 2021
- Athliini Smith & Evans, 2018
- Chasmatopterini Lacordaire, 1855
- Chnaunanthini Evans & Smith, 2020
- Colymbomorphini Blanchard, 1850
- Comophini Britton, 1978
- Dichelonychini Kirby, 1837
- Diphycerini Medvedev, 1952
- Diplotaxini Kirby, 1837
- Euchirini Hope, 1840
- Hopliini Latreille, 1829
- Langbianellini Prokofiev, 2015
- Macrodactylini Kirby, 1837
- Melolonthini Leach, 1819
- Pachypodini Erichson, 1840
- Pachytrichini Burmeister, 1855
- Phobetusini Evans & Smith, 2020
- Phyllotocidiini Britton, 1957
- Rhizotrogini Burmeister, 1855
- Systellopini Dalla Torre, 1912
- Warwickiini Evans & Smith, 2020
- †Cretomelolonthini Nikolajev, 1998
Subfamily Oncerinae LeConte, 1861 edit
Subfamily Orphninae Erichson, 1847 edit
Subfamily Pachydeminae Reitter, 1902 edit
- Aegosthetini Lacroix, 2007
- Pachydemini Burmeister, 1855
Subfamily Phaenomeridinae Erichson, 1847 edit
Subfamily Podolasiinae Howden, 1997 edit
Subfamily Rutelinae MacLeay, 1819 edit
- Adoretini Burmeister, 1844
- Alvarengiini Frey, 1975
- Anatistini Lacordaire, 1855
- Anomalini Streubel, 1839
- Anoplognathini MacLeay, 1819
- Geniatini Burmeister, 1844
- Rutelini MacLeay, 1819
Subfamily Scarabaeinae Latreille, 1802 edit
- Ateuchini Perty, 1830
- Byrrhidiini Davis, Deschodt & Scholtz, 2019
- Coprini Leach, 1815
- Deltochilini Lacordaire, 1855
- Dichotomiini Pereira, 1954
- Endroedyolini Davis, Deschodt & Scholtz, 2019
- Epactoidini Rossini et al, 2022
- Epilissini Lansberge, 1874
- Epirinini Lansberge, 1874
- Eucraniini Burmeister, 1873
- Gymnopleurini Streubel, 1846
- Odontolomini Davis, Deschodt & Scholtz, 2019
- Oniticellini Kolbe, 1905
- Onitini Castelnau, 1840
- Onthophagini Streubel, 1846
- Parachoriini Tarasov, 2017
- Phanaeini Hope, 1838
- Scarabaeini Latreille, 1802
- Sisyphini Mulsant, 1842
Subfamily Sericinae Kirby, 1837 edit
- Ablaberini Blanchard, 1850
- Diphucephalini Castelnau, 1840
- Sericini Kirby, 1837
Subfamily Sericoidinae Erichson, 1847 edit
- Automoliini Britton, 1978
- Heteronychini Lacordaire, 1855
- Liparetrini Burmeister, 1855
- Maechidiini Burmeister, 1855
- Phyllotocini Burmeister, 1855
- Scitalini Britton, 1957
- Sericoidini Erichson, 1847
Subfamily Termitotroginae Wasmann, 1918 edit
Subfamily Trichiinae Kolbe, 1897 edit
- Trichiini Fleming, 1821
Subfamily Valginae Schenkling, 1922 edit
- Valgini Mulsant, 1842
Subfamily †Cretoscarabaeinae Nikolajev, 1995 edit
Subfamily †Lithoscarabaeinae Nikolajev, 1992 edit
Subfamily †Prototroginae Nikolajev, 2000 edit
- †Prototrox Nikolajev, 2000
References edit
- ^ a b "Scarabaeidae Latreille, 1802". Catalogue of Life. Retrieved 2023-09-22.
- ^ Dietz, Lars; Seidel, Matthias; Eberle, Jonas; Misof, Bernhard; et al. (2023). "A transcriptome-based phylogeny of Scarabaeoidea confirms the sister group relationship of dung beetles and phytophagous pleurostict scarabs (Coleoptera)". Systematic Entomology. 48 (4). doi:10.1111/syen.12602.
- ^ Mondaca, José (2023). "A checklist of the Scarabaeoidea (Coleoptera) of Chile with exemplar live-photographed". Zootaxa. 5285 (2). doi:10.11646/ZOOTAXA.5285.2.1.